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RAF Tornado pilots had "struggled" to find targets until now, reports Jonathan Beale
RAF jets fired four missiles at Islamic State (IS) vehicles in Iraq overnight, the Ministry of Defence (MoD) has said.
It said two Tornados had fired missiles at an armed pick-up truck and transport vehicle west of Baghdad, adding that the strikes had been "successful".
Prime Minister David Cameron said there was no "walk-on-by option" for the UK when it came to dealing with IS.
Britons who join militants in Syria and Iraq are enemies of the UK and "should expect to be treated as such", he said.
"To those who have had all the advantages of being brought up in Britain, but who want to go and fight, let me say this - if you try and travel to Syria or Iraq, we will use everything at our disposal to stop you. Taking away your passport, prosecuting, convicting, imprisoning.
"And even if you're there already, we may even prevent you from coming back. You have declared your allegiance. You are an enemy of the UK and you should expect to be treated as such," Mr Cameron told the Conservative party conference.
'Command and control'His speech comes after RAF jets took part in further strikes in Iraq.
Commenting on the latest sorties, the MoD said the jets had been sent to examine a suspected command and control position of IS, the militant group also known as Isis or Isil.
"At the scene they were able to identify Isil activity and two vehicles, one of which was an armed pick-up truck," it said.
"Four Brimstone missiles were used to conduct a precision attack on the vehicles. Initial analysis indicates that the strikes were successful."
There has been no independent confirmation of the targets.
'We must respond'Defence Secretary Michael Fallon told the Tory conference in Birmingham that RAF planes had been conducting daily flights over Iraq since Parliament's vote.
He said "no responsible government" could ignore the challenge presented by IS.
"We didn't choose this fight, but for the security of our nation, we must respond," he said.
Foreign Secretary Philip Hammond told the conference that the "twisted ideology" of IS "can and must be driven out of Iraq".
"It is right that Britain shoulders its share of the burden in the great coalition of nations that has come together to defeat the menace of Isil, and we should be proud that we are doing so," he said.
He said the UK's strategy in the region was "comprehensive", including humanitarian aid, arming Kurdish fighters and cutting off IS funding.
About 40 nations have joined a coalition to tackle the militant group, which has seized large parts of Iraq and Syria in the past few months.
UK forces have now directly engaged IS twice since Parliament authorised involvement in the international military campaign last week.
On Tuesday, the MoD said Tornados had been on an "armed reconnaissance mission" when they were asked to help Kurdish troops in north-west Iraq.
The jets identified an IS heavy weapon position, which the MoD said was firing on Kurdish forces.
A guided bomb was used to attack the position, while a missile targeted an armed pick-up truck in the same area, it said.
Appeal for mercyThere were also reports of heavy fighting on Tuesday across a key border crossing between Iraq and Syria involving Kurdish forces and IS militants.
IS, which has established a self-declared "caliphate" in the territory it controls, is also holding a number of hostages.
Last month, in a video showing the beheading of British hostage David Haines the militants said the next to be killed may be Alan Henning - a taxi driver from Eccles who had been delivering aid to Syria when he was captured last year.
Mr Henning's wife, Barbara, has again appealed to IS leaders to show mercy.
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